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The 10.5.5 Update is recommended for all users running Mac OS X Leopard and includes general operating system fixes that enhance the stability, compatibility and security of your Mac. .
For more details about the update : http://support.apple.com/kb/HT2405?viewlocale=en_US .
For more details about the security update: http://support.apple.com/kb/HT1222?viewlocale=en_US .